Description
Key Characters:
Growth Form:
Annual or perennial herbs or small subshrubs.
Stems:
Stems prostrate or erect, usually 4-angled, glabrous to hispid or scabrid.
Roots:
Leaves:
Leaves simple.
Opposite or pseudowhorled.
Blades usually lanceolate to ovate or elliptic.
Margins with 1 to numerous filiform setae.
Sessile or petiolate.
Stipules adnate to petioles, forming a sheath.
Flowers:
Flowers in many-flowered clusters, usually axillary or terminal, subtended by 1-2 or more pairs of bract-like leaves, stipule-derived bracteoles with setae; hypanthium obovoid, turbinate, or obconical.
Flowers insect-pollinated, usually small, bisexual (perfect), sessile, often heterostylous.
Calyx lobes 2-4(-8) with open aestivation, usually deltate, oblong, or lanceolate, usually persistent, sometimes with intermediate teeth.
Corolla funnelform or salverform, the tube usually very slender, the lobes (3)4, spreading, valvate in bud; nectary disk usually present.
Stamens 4, alternate with the corolla lobes, inserted on corolla tube or throat, usually exserted; anthers dithecal, opening by longitudinal slits.
Ovary inferior, occasionally partly so, or very rarely superior, 2-celled; ovules 1 per cell, attached to middle of septum; style many as carpels, terminal, slender, filiform, exserted; stigmas lobed or capitate, dry or occasionally wet.
Fruit:
Fruit either a 2-valved capsule dehiscing from apex downward and the septum disappearing; or of 2 dimorphic mericarps; one dehiscent; the other indehiscent.
Seeds 2; usually glossy brown; oblong; ellipsoid; or ovoid; usually reticulate; ventrally grooved; with well-developed oily endosperm; or endosperm occasionally scanty or absent; with well-developed oily endosperm; or endosperm occasionally scanty or absent.
Ploidy:
Habitat:
Elevation Range: